Individual groups make the call.
Operationally availability depends how groups collate, for instance Yorkshire have an online system for responders to indicate their availability for up to 2 weeks (just upped from 1 week) ahead, other groups use sms responder (or similar) and ask at the point of need.
If groups were to extend social and event support type invites to neighbouring groups then the non callout arrangements would not require membership of more than one group and operationally mutual support arrangements could cover responders living in border areas (YR does this with Mitts for a NHS trust that straddles the border).
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